The offense welcomes back five starters but lost three-year starting QB Austen Arnaud, who finished his run as the second-leading passer in program history. In fall camp, juniors Jerome Tiller and Steele Jantz, a transfer from City College of San Francisco, will battle it out for the No. 1 job. Tiller was listed No. 1 on the depth chart both pre- and post-spring camp. ISU also lost a three-year starting running back in Alexander Robinson, who finished with 3,309 career yards, fourth-most in school history. The main to replace him will be sophomore Shontrelle Johnson, who showed signs of stardom in limited duty last year. The receiving corps is largely unproven, though veterans like Darius Darks, Darius Reynolds and Josh Lenz have shown flashes over the last few years.

On defense, seven starters are expected back. That unit was awful in rushing the passer last year with just 11 sacks, which ranked last in the Big 12 and 116th of 120 FBS teams. The strength of the defense will be at linebacker with juniors Jake Knott and A.J. Klein. Knott was a second-team All-Big 12 selection after finishing second in the league with 130 tackles. He also had a team-high four interceptions, four forced fumbles, a sack and a fumble recovery. Klein racked up 111 tackles, eight stops for loss and three interceptions.

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